No daughter colonies of the 4a were tested yet.
3 tested queens were mated at the mating station:
DE-2-501-91998-1991 DE-2-501-91999-1991 DE-2-503-91007-1991
6 daughter colonies of queens mated at the mating station were tested:
DE-2-503-6-1993 DE-2-503-39-1993 DE-2-503-67-1993 DE-2-503-78-1993 DE-2-503-89-1993 DE-2-503-100-1993