7 daughter colonies of the 4a were tested:
AT-99-117-3225-2016 AT-99-117-3226-2016 AT-99-117-3227-2016 AT-99-120-71208-2017 AT-99-120-71521-2017 AT-99-120-71522-2017 AT-99-120-71627-2016
19 tested queens were mated at the mating station:
AT-2-21-203-2017 AT-99-117-112-2017 AT-99-117-116-2017 AT-99-117-235-2017 AT-99-117-239-2017 AT-99-117-240-2017 AT-99-117-241-2017 AT-99-117-391-2017 AT-99-117-392-2017 AT-99-117-393-2017 AT-99-228-2-2017 AT-99-228-4-2017 AT-99-228-6-2017 AT-99-228-8-2017 AT-99-228-17-2017 AT-99-228-21-2017 AT-99-228-22-2017 AT-99-228-38-2017 AT-99-694-33-2017
15 daughter colonies of queens mated at the mating station were tested:
AT-99-228-1-2019 AT-99-228-4-2019 AT-99-228-60-2019 AT-99-228-65-2019 AT-99-228-67-2019 AT-99-228-68-2019 AT-99-228-69-2019 AT-99-228-75-2019 AT-99-228-77-2019 AT-99-228-78-2019 AT-99-228-81-2019 AT-99-228-83-2019 AT-99-228-84-2019 AT-99-228-85-2019 AT-99-228-98-2019