20 daughter colonies of the 4a were tested:
DE-2-322-161-2016 DE-6-1-82-2015 DE-6-1-84-2015 DE-6-1-85-2015 DE-6-1-86-2015 DE-6-1-87-2015 DE-6-1-88-2015 DE-6-1-91-2015 DE-6-1-92-2015 DE-6-1-95-2015 DE-6-1-98-2015 DE-6-1-99-2015 DE-6-1-100-2015 DE-6-1-103-2015 DE-6-1-104-2015 DE-6-1-105-2015 DE-6-1-106-2015 DE-6-1-108-2015 DE-6-1-110-2015 DE-6-1-111-2015
82 tested queens were mated at the mating station:
DE-6-1-179-2016 DE-6-1-181-2016 DE-6-1-187-2016 DE-6-1-196-2016 DE-6-1-200-2016 DE-6-1-201-2016 DE-6-1-210-2016 DE-6-1-249-2016 DE-6-1-316-2016 DE-6-1-318-2016 DE-6-1-320-2016 DE-6-1-325-2016 DE-6-1-327-2016 DE-6-1-328-2016 DE-6-1-329-2016 DE-6-1-330-2016 DE-6-1-332-2016 DE-6-1-333-2016 DE-6-1-529-2016 DE-6-1-532-2016 DE-6-1-536-2016 DE-6-1-538-2016 DE-6-1-539-2016 DE-6-1-540-2016 DE-6-1-542-2016 DE-6-1-547-2016 DE-6-1-551-2016 DE-6-1-568-2016 DE-6-1-582-2016 DE-6-1-585-2016 DE-6-1-596-2016 DE-6-1-603-2016 DE-6-1-608-2016 DE-6-1-613-2016 DE-6-1-628-2016 DE-6-1-636-2016 DE-6-1-637-2016 DE-6-1-640-2016 DE-6-1-649-2016 DE-6-1-678-2016 DE-6-1-679-2016 DE-6-1-690-2016 DE-6-1-692-2016 DE-6-1-714-2016 DE-6-1-717-2016 DE-6-1-720-2016 DE-6-1-723-2016 DE-6-1-727-2016 DE-6-1-734-2016 DE-6-1-737-2016 DE-6-1-740-2016 DE-6-1-743-2016 DE-6-1-746-2016 DE-6-1-770-2016 DE-6-1-771-2016 DE-6-1-772-2016 DE-6-1-1083-2016 DE-6-1-1105-2016 DE-6-1-1138-2016 DE-6-1-1141-2016 DE-6-1-1145-2016 DE-6-1-1199-2016 DE-6-1-1203-2016 DE-6-1-2004-2016 DE-6-1-2010-2016 DE-6-1-2016-2016 DE-6-1-2017-2016 DE-6-1-2021-2016 DE-6-1-2022-2016 DE-6-1-2024-2016 DE-6-1-2027-2016 DE-6-1-2028-2016 DE-6-1-2033-2016 DE-6-1-2040-2016 DE-6-1-2041-2016 DE-6-1-2046-2016 DE-6-1-2051-2016 DE-6-1-2052-2016 DE-6-1-2055-2016 DE-6-1-2056-2016 DE-6-1-2058-2016 DE-6-1-2064-2016
85 daughter colonies of queens mated at the mating station were tested:
DE-6-1-1-2018 DE-6-1-2-2018 DE-6-1-82-2018 DE-6-1-97-2018 DE-6-1-99-2018 DE-6-1-100-2018 DE-6-1-109-2018 DE-6-1-111-2018 DE-6-1-114-2018 DE-6-1-116-2018 DE-6-1-118-2018 DE-6-1-124-2018 DE-6-1-125-2018 DE-6-1-126-2018 DE-6-1-140-2018 DE-6-1-141-2018 DE-6-1-157-2018 DE-6-1-158-2018 DE-6-1-159-2018 DE-6-1-160-2018 DE-6-1-250-2018 DE-6-1-882-2018 DE-6-1-888-2018 DE-6-1-898-2018 DE-6-1-912-2018 DE-6-1-915-2018 DE-6-1-920-2018 DE-6-1-937-2018 DE-6-1-943-2018 DE-6-1-945-2018 DE-6-1-949-2018 DE-6-1-951-2018 DE-6-1-964-2018 DE-6-1-965-2018 DE-6-1-967-2018 DE-6-1-968-2018 DE-6-1-970-2018 DE-6-1-1004-2018 DE-6-1-1671-2018 DE-6-1-1676-2018 DE-6-1-1677-2018 DE-6-1-1679-2018 DE-6-1-1684-2018 DE-6-1-1685-2018 DE-6-1-1686-2018 DE-6-1-1715-2018 DE-6-1-1717-2018 DE-6-1-1722-2018 DE-6-1-1733-2018 DE-6-1-1909-2018 DE-6-1-1932-2018 DE-6-1-1933-2018 DE-6-1-1934-2018 DE-6-1-1935-2018 DE-6-1-1938-2018 DE-6-1-1944-2018 DE-6-1-1946-2018 DE-6-1-1947-2018 DE-6-1-1949-2018 DE-6-1-2017-2018 DE-6-1-2092-2018 DE-6-1-2101-2018 DE-6-1-2165-2018 DE-6-1-2167-2018 DE-6-1-2510-2018 DE-6-1-2515-2018 DE-6-1-2560-2018 DE-6-1-2596-2018 DE-6-1-2737-2019 DE-6-1-2817-2018 DE-6-1-2818-2018 DE-6-1-2820-2018 DE-18-51-1-2018 DE-18-51-2-2018 DE-18-51-3-2018 DE-18-51-4-2018 DE-18-51-5-2018 DE-18-51-6-2018 DE-18-51-7-2018 DE-18-51-8-2018 DE-18-51-9-2018 DE-18-51-10-2018 DE-18-51-11-2018 DE-18-51-12-2018 DE-18-51-1725-2018