7 daughter colonies of the 4a were tested:
DE-6-1-544-2000 DE-6-1-549-2000 DE-6-1-550-2000 DE-6-1-551-2000 DE-6-1-552-2000 DE-6-1-556-2000 DE-6-1-557-2000
70 tested queens were mated at the mating station:
DE-6-1-1-2001 DE-6-1-3-2001 DE-6-1-7-2001 DE-6-1-8-2001 DE-6-1-9-2001 DE-6-1-10-2001 DE-6-1-13-2001 DE-6-1-14-2001 DE-6-1-15-2001 DE-6-1-19-2001 DE-6-1-20-2001 DE-6-1-21-2001 DE-6-1-22-2001 DE-6-1-24-2001 DE-6-1-25-2001 DE-6-1-26-2001 DE-6-1-27-2001 DE-6-1-29-2001 DE-6-1-31-2001 DE-6-1-32-2001 DE-6-1-35-2001 DE-6-1-36-2001 DE-6-1-37-2001 DE-6-1-38-2001 DE-6-1-41-2001 DE-6-1-48-2001 DE-6-1-49-2001 DE-6-1-50-2001 DE-6-1-51-2001 DE-6-1-58-2001 DE-6-1-60-2001 DE-6-1-62-2001 DE-6-1-63-2001 DE-6-1-67-2001 DE-6-1-85-2001 DE-6-1-695-2001 DE-6-1-700-2001 DE-6-1-701-2001 DE-6-1-707-2001 DE-6-1-709-2001 DE-6-1-711-2001 DE-6-1-712-2001 DE-6-1-713-2001 DE-6-1-717-2001 DE-6-1-719-2001 DE-6-1-1280-2001 DE-6-1-1289-2001 DE-6-1-1290-2001 DE-6-1-1293-2001 DE-6-1-1298-2001 DE-6-1-1308-2001 DE-6-1-1312-2001 DE-6-1-1314-2001 DE-6-1-1320-2001 DE-6-1-1321-2001 DE-6-1-1323-2001 DE-6-1-1326-2001 DE-6-1-1352-2001 DE-6-1-1353-2001 DE-6-1-1355-2001 DE-6-1-1358-2001 DE-6-1-1391-2001 DE-6-1-1392-2001 DE-6-1-1393-2001 DE-6-1-1400-2001 DE-6-1-1412-2001 DE-6-1-1423-2001 DE-6-1-1433-2001 DE-6-1-1441-2001 DE-6-1-1495-2001
57 daughter colonies of queens mated at the mating station were tested:
DE-6-1-315-2004 DE-6-1-320-2004 DE-6-1-323-2004 DE-6-1-330-2004 DE-6-1-332-2004 DE-6-1-371-2004 DE-6-1-381-2004 DE-6-1-388-2004 DE-6-1-393-2004 DE-6-1-400-2004 DE-6-1-406-2004 DE-6-1-411-2004 DE-6-1-427-2004 DE-6-1-435-2004 DE-6-1-472-2004 DE-6-1-481-2004 DE-6-1-488-2004 DE-6-1-493-2004 DE-6-1-494-2004 DE-6-1-1167-2003 DE-6-1-1168-2003 DE-6-1-1169-2003 DE-6-1-1170-2003 DE-6-1-1171-2003 DE-6-1-1174-2003 DE-6-1-1175-2003 DE-6-1-1177-2003 DE-6-1-1538-2004 DE-6-1-1541-2004 DE-6-1-1547-2004 DE-6-1-1559-2004 DE-6-1-1563-2004 DE-6-1-1565-2004 DE-6-1-1566-2004 DE-6-1-1568-2004 DE-6-1-1570-2004 DE-6-1-1573-2004 DE-6-1-1574-2004 DE-6-1-1576-2004 DE-6-1-1578-2004 DE-6-1-1581-2004 DE-6-1-1595-2004 DE-6-1-1599-2004 DE-6-1-1606-2004 DE-6-1-1610-2004 DE-6-1-1613-2004 DE-6-1-1619-2004 DE-6-1-1622-2004 DE-6-1-1624-2004 DE-6-1-1629-2004 DE-6-1-1631-2004 DE-6-1-1632-2004 DE-6-1-1639-2004 DE-6-1-1641-2004 DE-6-1-1647-2004 DE-6-1-1648-2004 DE-6-1-1656-2004