17 daughter colonies of the 4a were tested:
AT-2-3-861-2018  AT-2-3-863-2018  AT-2-3-883-2018  AT-2-3-884-2018  AT-2-3-887-2018  AT-2-3-889-2018  AT-2-3-962-2019  AT-2-7-844-2018  AT-2-7-883-2018  AT-2-7-884-2018  AT-2-7-886-2018  AT-2-8-2929-2019  AT-2-8-2930-2019  AT-2-8-2931-2019  AT-2-8-2932-2019  AT-2-8-2933-2019  AT-2-8-2934-2019  
79  tested queens were mated at the mating station:
AT-2-3-901-2019  AT-2-3-902-2019  AT-2-3-904-2019  AT-2-3-905-2019  AT-2-3-906-2019  AT-2-3-907-2019  AT-2-3-913-2019  AT-2-3-918-2019  AT-2-3-922-2019  AT-2-3-931-2019  AT-2-3-934-2019  AT-2-3-943-2019  AT-2-3-944-2019  AT-2-3-945-2019  AT-2-3-956-2019  AT-2-3-962-2019  AT-2-6-670-2019  AT-2-7-902-2019  AT-2-7-903-2019  AT-2-7-916-2019  AT-2-7-917-2019  AT-2-7-918-2019  AT-2-7-919-2019  AT-2-7-920-2019  AT-2-7-927-2019  AT-2-7-933-2019  AT-2-7-934-2019  AT-2-7-938-2019  AT-2-7-947-2019  AT-2-7-9030-2019  AT-2-7-9035-2019  AT-2-7-9036-2019  AT-2-7-9037-2019  AT-2-8-1901-2019  AT-2-8-1902-2019  AT-2-8-1903-2019  AT-2-8-1904-2019  AT-2-8-1905-2019  AT-2-8-1906-2019  AT-2-8-1907-2019  AT-2-8-1915-2019  AT-2-8-1916-2019  AT-2-8-1917-2019  AT-2-8-1918-2019  AT-2-8-1919-2019  AT-2-8-1920-2019  AT-2-8-1922-2019  AT-2-8-1923-2019  AT-2-8-1924-2019  AT-2-8-2902-2019  AT-2-8-2903-2019  AT-2-8-2904-2019  AT-2-8-2905-2019  AT-2-8-2920-2019  AT-2-8-2921-2019  AT-2-8-2923-2019  AT-2-8-3901-2019  AT-2-8-3903-2019  AT-2-8-4901-2019  AT-2-8-4902-2019  AT-2-8-4903-2019  AT-2-8-4904-2019  AT-2-8-4905-2019  AT-2-8-4911-2019  AT-2-8-4912-2019  AT-99-364-20-2019  AT-99-364-21-2019  AT-99-364-22-2019  AT-99-364-23-2019  AT-99-364-24-2019  AT-99-364-25-2019  AT-99-364-26-2019  AT-99-364-27-2019  AT-99-495-103-2019  AT-99-495-104-2019  AT-99-495-105-2019  AT-99-495-106-2019  AT-99-495-107-2019  AT-99-495-108-2019  
183  daughter colonies of queens mated at the mating station were tested:
DE-7-215-85-2022  AT-2-1-63-2021  AT-2-1-130-2021  AT-2-1-138-2021  AT-2-1-139-2021  AT-2-1-184-2021  AT-2-1-207-2021  AT-2-1-215-2021  AT-2-1-221-2021  AT-2-1-230-2021  AT-2-1-232-2021  AT-2-1-239-2021  AT-2-1-264-2021  AT-2-1-342-2021  AT-2-1-348-2021  AT-2-1-349-2021  AT-2-1-390-2021  AT-2-1-401-2021  AT-2-1-603-2021  AT-2-1-614-2021  AT-2-1-615-2021  AT-2-1-621-2021  AT-2-1-671-2021  AT-2-1-775-2021  AT-2-1-823-2021  AT-2-1-906-2021  AT-2-1-1018-2021  AT-2-1-1030-2021  AT-2-1-1045-2021  AT-2-1-1091-2021  AT-2-1-1118-2021  AT-2-1-1121-2021  AT-2-1-1272-2021  AT-2-1-1291-2021  AT-2-1-1299-2021  AT-2-1-1390-2021  AT-2-1-1392-2021  AT-2-1-1445-2021  AT-2-1-1483-2021  AT-2-1-1486-2021  AT-2-1-1489-2021  AT-2-1-1539-2021  AT-2-1-1569-2021  AT-2-1-1574-2021  AT-2-1-1577-2021  AT-2-1-1578-2021  AT-2-1-2461-2021  AT-2-1-2466-2021  AT-2-1-2753-2021  AT-2-7-104-2021  AT-2-7-106-2021  AT-2-7-107-2021  AT-2-7-110-2021  AT-2-7-111-2021  AT-2-7-112-2021  AT-2-7-114-2021  AT-2-7-115-2021  AT-2-7-116-2021  AT-2-7-119-2021  AT-2-7-121-2021  AT-2-7-125-2021  AT-2-7-128-2021  AT-2-7-129-2021  AT-2-7-133-2021  AT-2-7-134-2021  AT-2-7-135-2021  AT-2-7-142-2021  AT-2-7-147-2021  AT-2-7-148-2021  AT-2-7-151-2021  AT-2-7-153-2021  AT-2-7-156-2021  AT-2-7-157-2021  AT-2-7-162-2021  AT-2-7-163-2021  AT-2-7-165-2021  AT-2-7-172-2021  AT-2-7-173-2021  AT-2-7-174-2021  AT-2-7-176-2021  AT-2-7-178-2021  AT-2-7-180-2021  AT-2-7-181-2021  AT-2-7-184-2021  AT-2-7-189-2021  AT-2-7-191-2021  AT-2-7-192-2021  AT-2-7-195-2021  AT-2-7-1026-2021  AT-2-7-1029-2021  AT-2-7-1034-2021  AT-2-7-1035-2021  AT-2-7-1036-2021  AT-2-7-1042-2021  AT-2-7-1043-2021  AT-2-7-1050-2021  AT-2-7-1051-2021  AT-2-7-1052-2021  AT-2-7-1054-2021  AT-2-7-1057-2021  AT-2-7-1059-2021  AT-2-7-1062-2021  AT-2-7-1067-2021  AT-2-7-1069-2021  AT-2-7-1070-2021  AT-2-7-1071-2021  AT-2-7-1073-2021  AT-2-7-1075-2021  AT-2-7-1077-2021  AT-2-7-1080-2021  AT-2-7-1082-2021  AT-2-7-1083-2021  AT-2-7-1084-2021  AT-2-7-1085-2021  AT-2-8-1101-2021  AT-2-8-1102-2021  AT-2-8-1104-2021  AT-2-8-1105-2021  AT-2-8-1106-2021  AT-2-8-1107-2021  AT-2-8-1108-2021  AT-2-8-1109-2021  AT-2-8-1110-2021  AT-2-8-1113-2021  AT-2-8-4001-2022  AT-2-8-4002-2022  AT-2-8-4003-2022  AT-2-8-4004-2022  AT-2-8-4005-2022  AT-2-8-4006-2022  AT-2-8-4007-2022  AT-2-8-4008-2022  AT-2-8-4111-2021  AT-2-8-4112-2021  AT-2-8-4113-2021  AT-2-8-4114-2021  AT-2-8-4115-2021  AT-2-8-4116-2021  AT-2-8-4117-2021  AT-2-8-4118-2021  AT-2-8-4119-2021  AT-2-8-4120-2021  AT-2-8-4122-2021  AT-99-266-2-2021  AT-99-266-3-2021  AT-99-266-5-2021  AT-99-266-8-2021  AT-99-266-12-2021  AT-99-266-17-2020  AT-99-266-18-2020  AT-99-266-19-2020  AT-99-266-19-2021  AT-99-266-20-2020  AT-99-266-21-2020  AT-99-266-22-2020  AT-99-266-22-2021  AT-99-266-23-2020  AT-99-266-24-2020  AT-99-266-25-2020  AT-99-266-25-2021  AT-99-266-26-2020  AT-99-266-26-2021  AT-99-266-27-2020  AT-99-266-27-2021  AT-99-266-28-2020  AT-99-266-41-2021  AT-99-266-42-2021  AT-99-266-43-2021  AT-99-266-44-2021  AT-99-266-45-2021  AT-99-266-46-2021  AT-99-364-21-2021  AT-99-364-22-2021  AT-99-364-23-2021  AT-99-364-24-2021  AT-99-364-25-2021  AT-99-364-28-2021  AT-99-364-29-2021  AT-99-364-30-2021  AT-99-364-31-2021  AT-99-364-122-2021  AT-99-364-123-2021  AT-99-364-124-2021