9 daughter colonies of the 4a were tested:
AT-99-106-21932-1992 AT-99-106-22155-1992 AT-99-106-23443-1992 AT-99-106-23546-1992 AT-99-106-23593-1992 AT-99-106-26962-1991 AT-99-110-17545-1991 AT-99-110-22155-1992 AT-99-110-22782-1992
212 tested queens were mated at the mating station:
AT-2-2-15378-1993 AT-2-2-15398-1993 AT-2-2-15448-1993 AT-2-2-15503-1993 AT-2-2-15531-1993 AT-2-2-15544-1993 AT-2-2-15566-1993 AT-2-2-15580-1993 AT-2-2-15597-1993 AT-2-2-15618-1993 AT-2-2-15631-1993 AT-2-2-15683-1993 AT-2-2-15708-1993 AT-2-2-15729-1993 AT-2-2-15752-1993 AT-2-2-15780-1993 AT-2-2-15807-1993 AT-2-2-15838-1993 AT-2-2-15861-1993 AT-2-2-15929-1993 AT-2-2-15968-1993 AT-2-2-15993-1993 AT-2-2-16016-1993 AT-2-2-16029-1993 AT-2-2-16052-1993 AT-2-2-16102-1993 AT-2-2-16129-1993 AT-2-2-16152-1993 AT-2-2-16171-1993 AT-2-2-16215-1993 AT-2-2-16235-1993 AT-2-2-16255-1993 AT-2-2-16357-1993 AT-2-2-16387-1993 AT-2-2-16411-1993 AT-2-2-16426-1993 AT-2-2-16443-1993 AT-2-2-16459-1993 AT-2-2-16492-1993 AT-2-2-16508-1993 AT-2-2-16574-1993 AT-2-2-16595-1993 AT-2-2-16606-1993 AT-2-2-16659-1993 AT-2-2-16672-1993 AT-2-2-16692-1993 AT-2-2-16720-1993 AT-2-2-16735-1993 AT-2-2-16747-1993 AT-2-2-16761-1993 AT-2-2-16795-1993 AT-2-2-16822-1993 AT-2-2-16836-1993 AT-2-2-16858-1993 AT-2-2-16876-1993 AT-2-2-16916-1993 AT-2-2-16957-1993 AT-2-2-16965-1993 AT-2-2-16984-1993 AT-2-2-16994-1993 AT-2-2-17011-1993 AT-2-2-17022-1993 AT-2-2-17034-1993 AT-2-2-17049-1993 AT-2-2-17063-1993 AT-2-2-17077-1993 AT-2-2-17088-1993 AT-2-2-17105-1993 AT-2-2-17116-1993 AT-2-2-17140-1993 AT-2-2-17169-1993 AT-2-2-17192-1993 AT-2-2-17207-1993 AT-2-2-17227-1993 AT-2-2-17261-1993 AT-2-2-17297-1993 AT-2-2-17326-1993 AT-2-2-17352-1993 AT-2-2-17380-1993 AT-2-2-17402-1993 AT-2-2-17422-1993 AT-2-2-17436-1993 AT-2-2-17450-1993 AT-2-2-17473-1993 AT-2-2-17497-1993 AT-2-2-17510-1993 AT-2-2-17555-1993 AT-2-2-17608-1993 AT-2-2-17626-1993 AT-2-2-17644-1993 AT-2-2-17655-1993 AT-2-2-17671-1993 AT-2-2-17706-1993 AT-2-2-17717-1993 AT-2-2-17734-1993 AT-2-2-17749-1993 AT-2-2-17772-1993 AT-2-2-17786-1993 AT-2-2-17797-1993 AT-2-2-17814-1993 AT-2-2-17831-1993 AT-2-2-17844-1993 AT-2-2-17856-1993 AT-2-2-17889-1993 AT-2-2-17907-1993 AT-2-2-17933-1993 AT-2-2-17954-1993 AT-2-2-17968-1993 AT-2-2-17981-1993 AT-2-2-18001-1993 AT-2-2-18028-1993 AT-2-2-18044-1993 AT-2-2-18058-1993 AT-2-2-18070-1993 AT-2-2-18083-1993 AT-2-2-18098-1993 AT-2-2-18116-1993 AT-2-2-18127-1993 AT-2-2-18164-1993 AT-2-2-18191-1993 AT-2-2-18238-1993 AT-2-2-18270-1993 AT-2-2-18300-1993 AT-2-2-18315-1993 AT-2-2-18330-1993 AT-2-2-18353-1993 AT-2-2-18399-1993 AT-2-2-18413-1993 AT-2-2-18457-1993 AT-2-2-18469-1993 AT-2-2-18479-1993 AT-2-2-18500-1993 AT-2-2-18522-1993 AT-2-2-18535-1993 AT-2-2-18543-1993 AT-2-2-18553-1993 AT-2-2-18570-1993 AT-2-2-18582-1993 AT-2-2-18594-1993 AT-2-2-18615-1993 AT-2-2-18633-1993 AT-2-2-18646-1993 AT-2-2-18658-1993 AT-2-2-18666-1993 AT-2-2-18677-1993 AT-2-2-18695-1993 AT-2-2-18709-1993 AT-2-2-18745-1993 AT-2-2-18793-1993 AT-2-2-18810-1993 AT-2-2-18865-1993 AT-2-2-18914-1993 AT-2-2-18974-1993 AT-2-2-18990-1993 AT-2-2-19015-1993 AT-2-2-19069-1993 AT-2-2-19095-1993 AT-2-2-19107-1993 AT-2-2-19157-1993 AT-2-2-19181-1993 AT-2-2-19204-1993 AT-99-106-11303-1993 AT-99-106-11390-1993 AT-99-106-11398-1993 AT-99-106-11424-1993 AT-99-106-11490-1993 AT-99-106-11495-1993 AT-99-106-11498-1993 AT-99-106-11501-1993 AT-99-106-11513-1993 AT-99-106-11521-1993 AT-99-106-11536-1993 AT-99-106-11541-1993 AT-99-106-12219-1993 AT-99-106-12781-1993 AT-99-106-13139-1993 AT-99-106-13159-1993 AT-99-106-13383-1993 AT-99-106-13924-1993 AT-99-106-13954-1993 AT-99-106-13973-1993 AT-99-106-14025-1993 AT-99-106-14555-1993 AT-99-106-14594-1993 AT-99-106-14666-1993 AT-99-106-14713-1993 AT-99-106-14782-1993 AT-99-106-14816-1993 AT-99-106-14842-1993 AT-99-106-14869-1993 AT-99-106-14891-1993 AT-99-106-18076-1993 AT-99-106-18563-1993 AT-99-106-19218-1993 AT-99-106-19575-1993 AT-99-106-21261-1993 AT-99-106-21926-1993 AT-99-106-22389-1993 AT-99-106-22799-1993 AT-99-106-22852-1993 AT-99-106-23893-1993 AT-99-106-25937-1993 AT-99-119-15314-1993 AT-99-119-15319-1993 AT-99-119-15320-1993 AT-99-119-15325-1993 AT-99-119-15327-1993 AT-99-119-15328-1993 AT-99-119-15343-1993 AT-99-119-15345-1993 AT-99-119-15348-1993 AT-99-119-15506-1993
229 daughter colonies of queens mated at the mating station were tested:
AT-2-2-10410-1996 AT-2-2-10435-1996 AT-2-2-10460-1996 AT-2-2-16859-1995 AT-2-2-17117-1995 AT-2-2-17353-1995 AT-2-2-17451-1995 AT-2-2-17578-1995 AT-2-2-18060-1995 AT-2-2-18085-1995 AT-2-2-18099-1995 AT-2-2-18117-1995 AT-2-2-18471-1995 AT-2-2-19134-1995 AT-2-2-19182-1995 AT-2-2-19205-1995 AT-2-2-19227-1995 AT-2-2-19242-1995 AT-2-2-19279-1995 AT-2-2-19293-1995 AT-2-2-19321-1995 AT-2-2-19348-1995 AT-2-2-19369-1995 AT-2-2-19384-1995 AT-2-2-19417-1995 AT-2-2-19478-1995 AT-2-2-19500-1995 AT-2-2-19515-1995 AT-2-2-19545-1995 AT-2-2-19584-1995 AT-2-2-19652-1995 AT-2-2-19680-1995 AT-2-2-19693-1995 AT-2-2-19732-1995 AT-2-2-19747-1995 AT-2-2-19768-1995 AT-2-2-19818-1995 AT-2-2-19826-1995 AT-2-2-19841-1995 AT-2-2-20116-1995 AT-2-2-20129-1995 AT-2-2-20142-1995 AT-2-2-20162-1995 AT-2-2-20175-1995 AT-2-2-20186-1995 AT-2-2-20199-1995 AT-2-2-20220-1995 AT-2-2-20234-1995 AT-2-2-20257-1995 AT-2-2-20276-1995 AT-2-2-20283-1995 AT-2-2-20336-1995 AT-2-2-20483-1994 AT-2-2-20488-1994 AT-2-2-20492-1994 AT-2-2-20495-1994 AT-2-2-20499-1994 AT-2-2-20506-1994 AT-2-2-20517-1994 AT-2-2-20518-1994 AT-2-2-20531-1994 AT-2-2-20553-1994 AT-2-2-20563-1995 AT-2-2-20566-1994 AT-2-2-20571-1994 AT-2-2-20578-1994 AT-2-2-20580-1994 AT-2-2-20587-1994 AT-2-2-20589-1994 AT-2-2-20591-1994 AT-2-2-20597-1994 AT-2-2-20599-1994 AT-2-2-20602-1995 AT-2-2-20609-1994 AT-2-2-20612-1994 AT-2-2-20643-1994 AT-2-2-20648-1995 AT-2-2-20649-1994 AT-2-2-20652-1994 AT-2-2-20653-1994 AT-2-2-20656-1994 AT-2-2-20658-1994 AT-2-2-20662-1994 AT-2-2-20664-1994 AT-2-2-20666-1995 AT-2-2-20667-1994 AT-2-2-20670-1994 AT-2-2-20672-1994 AT-2-2-20675-1994 AT-2-2-20677-1994 AT-2-2-20678-1994 AT-2-2-20680-1994 AT-2-2-20685-1995 AT-2-2-20689-1994 AT-2-2-20690-1994 AT-2-2-20697-1994 AT-2-2-20700-1994 AT-2-2-20702-1994 AT-2-2-20705-1994 AT-2-2-20708-1994 AT-2-2-20712-1995 AT-2-2-20720-1994 AT-2-2-20723-1994 AT-2-2-20728-1994 AT-2-2-20731-1994 AT-2-2-20738-1994 AT-2-2-20762-1994 AT-2-2-20766-1994 AT-2-2-20768-1994 AT-2-2-20773-1994 AT-2-2-20784-1994 AT-2-2-20802-1994 AT-2-2-20809-1994 AT-2-2-20849-1994 AT-2-2-20850-1994 AT-2-2-20855-1994 AT-2-2-20856-1994 AT-2-2-20859-1994 AT-2-2-20884-1994 AT-2-2-20889-1994 AT-2-2-20900-1994 AT-2-2-20901-1994 AT-2-2-20903-1994 AT-2-2-20904-1994 AT-2-2-20909-1994 AT-2-2-20913-1994 AT-2-2-20915-1994 AT-2-2-20916-1994 AT-2-2-20919-1994 AT-2-2-20927-1994 AT-2-2-20931-1994 AT-2-2-20938-1994 AT-2-2-20939-1994 AT-2-2-21859-1995 AT-2-2-21875-1995 AT-99-106-10625-1995 AT-99-106-10783-1995 AT-99-106-10852-1995 AT-99-106-10891-1995 AT-99-106-10910-1995 AT-99-106-10947-1995 AT-99-106-10998-1995 AT-99-106-11004-1995 AT-99-106-11065-1995 AT-99-106-11073-1995 AT-99-106-11113-1995 AT-99-106-11142-1995 AT-99-106-11199-1995 AT-99-106-11209-1995 AT-99-106-11377-1995 AT-99-106-11384-1995 AT-99-106-11387-1995 AT-99-106-11391-1995 AT-99-106-11402-1995 AT-99-106-11409-1995 AT-99-106-11433-1995 AT-99-106-11448-1995 AT-99-106-11452-1995 AT-99-106-11459-1995 AT-99-106-11582-1995 AT-99-106-11636-1995 AT-99-106-11689-1995 AT-99-106-11702-1995 AT-99-106-11919-1995 AT-99-106-11931-1995 AT-99-106-11952-1995 AT-99-106-11963-1995 AT-99-106-11967-1995 AT-99-106-13151-1995 AT-99-106-13592-1995 AT-99-106-17609-1994 AT-99-106-17627-1994 AT-99-106-17656-1994 AT-99-106-17672-1994 AT-99-106-17707-1994 AT-99-106-17718-1994 AT-99-106-17832-1994 AT-99-106-17870-1994 AT-99-106-17890-1994 AT-99-106-17908-1994 AT-99-106-17955-1994 AT-99-106-17982-1994 AT-99-106-18029-1994 AT-99-106-18071-1994 AT-99-106-18142-1994 AT-99-106-18165-1994 AT-99-106-18225-1994 AT-99-106-18255-1994 AT-99-106-18271-1994 AT-99-106-18301-1994 AT-99-106-18316-1994 AT-99-106-18634-1994 AT-99-106-18647-1994 AT-99-106-25894-1995 AT-99-106-25943-1995 AT-99-106-25956-1995 AT-99-106-27062-1995 AT-99-107-11651-1994 AT-99-119-10138-1996 AT-99-119-10150-1996 AT-99-119-10162-1996 AT-99-119-10170-1996 AT-99-119-10173-1996 AT-99-119-10175-1996 AT-99-119-10177-1996 AT-99-119-10180-1996 AT-99-119-11542-1994 AT-99-119-11548-1994 AT-99-119-11562-1994 AT-99-119-11761-1994 AT-99-119-11802-1994 AT-99-119-11804-1994 AT-99-119-23188-1995 AT-99-119-23190-1995 AT-99-127-22814-1994 AT-99-127-22831-1994 AT-99-127-22896-1994 AT-99-127-22915-1994 AT-99-127-22936-1994 AT-99-127-22957-1994 AT-99-127-22964-1994 AT-99-127-22992-1994 AT-99-127-23006-1994 AT-99-359-13800-1995 AT-99-359-16538-1995 AT-99-359-16541-1995 AT-99-359-16550-1995 AT-99-359-16556-1995 AT-99-359-16560-1995