5 daughter colonies of the 4a were tested:
AT-2-2-15838-1993 AT-2-2-16016-1993 AT-2-2-16152-1993 AT-2-2-16235-1993 AT-2-2-18974-1993
2 tested queens were mated at the mating station:
AT-2-2-20832-1994 AT-2-2-20838-1994
10 daughter colonies of queens mated at the mating station were tested:
AT-2-2-15438-1997 AT-2-2-15444-1997 AT-2-2-15452-1997 AT-2-2-15455-1997 AT-2-2-15462-1997 AT-2-2-15464-1997 AT-2-2-15467-1997 AT-2-2-15472-1997 AT-2-2-18572-1995 AT-2-2-18583-1995