9 daughter colonies of the 4a were tested:
AT-2-2-16261-1997 AT-2-2-16263-1997 AT-2-2-16273-1997 AT-2-2-16280-1997 AT-2-2-16287-1997 AT-2-2-16291-1997 AT-2-2-16292-1997 AT-2-2-16295-1997 AT-2-2-16301-1997
7 tested queens were mated at the mating station:
AT-2-2-10940-1998 AT-2-2-10941-1998 AT-2-2-10944-1998 AT-2-2-10945-1998 AT-2-2-10948-1998 AT-2-2-10949-1998 AT-2-2-10950-1998
20 daughter colonies of queens mated at the mating station were tested:
AT-2-2-22168-1999 AT-2-2-22171-1999 AT-2-2-22176-1999 AT-2-2-22177-1999 AT-2-2-22180-1999 AT-2-2-22186-1999 AT-2-2-22189-1999 AT-2-2-22200-1999 AT-2-2-22202-1999 AT-2-2-22381-1999 AT-2-2-22383-1999 AT-2-2-22386-1999 AT-2-2-22391-1999 AT-2-2-22400-1999 AT-2-2-22520-1999 AT-2-2-22526-1999 AT-2-2-22550-1999 AT-2-2-22553-1999 AT-2-2-22556-1999 AT-2-2-22561-1999