11 daughter colonies of the 4a were tested:
CH-51-100-1892-1985 CH-51-100-2214-1986 CH-51-100-2483-1985 CH-51-100-2484-1985 CH-51-101-2321-1987 CH-51-101-2322-1987 CH-51-101-2774-1988 CH-51-101-2775-1988 CH-51-101-2776-1988 CH-51-106-2091-1986 CH-51-106-2204-1986
25 tested queens were mated at the mating station:
CH-51-4-3432-1990 CH-51-4-3433-1990 CH-51-4-3796-1990 CH-51-100-3288-1990 CH-51-100-3289-1990 CH-51-102-3449-1990 CH-51-102-3451-1990 CH-51-102-3452-1990 CH-51-106-3510-1990 CH-51-106-3511-1990 CH-51-106-3513-1990 CH-51-106-3514-1990 CH-51-106-3515-1990 CH-51-109-3359-1990 CH-51-109-3360-1990 CH-51-109-3361-1990 CH-51-109-3362-1990 CH-51-109-3363-1990 CH-51-109-3364-1990 CH-51-109-3365-1990 CH-51-109-3366-1990 CH-51-109-3367-1990 CH-51-109-3368-1990 CH-51-109-3369-1990 CH-51-109-3370-1990
5 daughter colonies of queens mated at the mating station were tested:
CH-51-4-5328-1992 CH-51-102-3818-1991 CH-51-102-3819-1991 CH-51-102-3820-1991 CH-51-109-3991-1991