9 daughter colonies of the 4a were tested:
DE-13-4-1-1993 DE-13-4-5-1993 DE-13-4-8-1993 DE-13-4-10-1993 DE-13-4-12-1993 DE-13-4-13-1993 DE-13-4-22-1993 DE-13-4-23-1993 DE-13-4-26-1993
8 tested queens were mated at the mating station:
DE-13-4-5-1994 DE-13-4-10-1994 DE-13-4-11-1994 DE-13-4-17-1994 DE-13-4-19-1994 DE-13-4-26-1994 DE-13-4-35-1994 DE-13-4-36-1994
2 daughter colonies of queens mated at the mating station were tested:
DE-13-4-36-1996 DE-13-4-39-1996