2 daughter colonies of the 4a were tested:
DE-15-29-17-1988 DE-15-29-18-1988
5 tested queens were mated at the mating station:
DE-15-29-37-1989 DE-15-29-38-1989 DE-15-29-39-1989 DE-15-29-56-1989 DE-15-29-57-1989
1 daughter colonies of queens mated at the mating station were tested:
DE-15-29-96-1991