No daughter colonies of the 4a were tested yet.
40 tested queens were mated at the mating station:
DE-4-1-284-1989 DE-4-1-287-1989 DE-4-1-328-1989 DE-4-1-331-1989 DE-4-1-333-1989 DE-4-1-340-1989 DE-4-1-342-1989 DE-4-1-343-1989 DE-4-1-345-1989 DE-4-1-350-1989 DE-4-1-362-1989 DE-4-1-428-1989 DE-4-1-432-1989 DE-4-1-435-1989 DE-4-1-436-1989 DE-4-1-439-1989 DE-4-1-442-1989 DE-4-1-443-1989 DE-4-1-445-1989 DE-4-1-446-1989 DE-4-1-466-1989 DE-4-1-467-1989 DE-4-1-468-1989 DE-4-1-473-1989 DE-4-1-478-1989 DE-4-1-591-1989 DE-4-1-592-1989 DE-4-1-597-1989 DE-4-1-601-1989 DE-4-1-603-1989 DE-4-1-635-1989 DE-4-1-642-1989 DE-4-1-645-1989 DE-4-1-949-1989 DE-4-1-959-1989 DE-4-1-971-1989 DE-4-1-99462-1989 DE-4-1-99463-1989 DE-4-1-99469-1989 DE-4-1-99471-1989
16 daughter colonies of queens mated at the mating station were tested:
DE-4-9-2-1991 DE-4-9-6-1991 DE-4-9-12-1991 DE-4-9-119-1991 DE-4-9-120-1991 DE-4-9-122-1991 DE-4-9-123-1991 DE-4-9-124-1991 DE-4-9-125-1991 DE-4-9-128-1991 DE-4-9-130-1991 DE-4-9-131-1991 DE-4-9-132-1991 DE-4-9-133-1991 DE-4-9-136-1991 DE-4-9-137-1991