7 daughter colonies of the 4a were tested:
DE-4-5-1230-1986 DE-4-5-1231-1986 DE-4-5-1232-1986 DE-4-5-1233-1986 DE-4-5-1234-1986 DE-4-5-1235-1986 DE-4-5-1236-1986
14 tested queens were mated at the mating station:
DE-4-34-2-1985 DE-4-34-3-1985 DE-4-34-4-1985 DE-4-34-5-1985 DE-4-34-6-1985 DE-4-34-7-1985 DE-4-34-8-1985 DE-4-34-14-1985 DE-4-34-15-1985 DE-4-34-16-1985 DE-4-34-17-1985 DE-4-34-18-1985 DE-4-34-19-1985 DE-4-34-20-1985
17 daughter colonies of queens mated at the mating station were tested:
DE-4-34-11-1987 DE-4-34-12-1987 DE-4-34-13-1987 DE-4-34-14-1987 DE-4-34-15-1987 DE-4-34-16-1987 DE-4-34-33-1988 DE-4-34-34-1988 DE-4-34-36-1988 DE-4-34-37-1988 DE-4-34-42-1988 DE-4-34-46-1988 DE-4-34-92-1988 DE-4-34-96-1988 DE-4-34-99-1988 DE-4-34-100-1988 DE-4-34-108-1988