7 daughter colonies of the 4a were tested:
DE-4-5-1495-1991 DE-4-5-1496-1991 DE-4-5-1500-1991 DE-4-5-1504-1991 DE-4-5-1505-1991 DE-4-5-1506-1991 DE-4-5-1509-1991
24 tested queens were mated at the mating station:
DE-4-18-749-1988 DE-4-18-750-1988 DE-4-18-751-1988 DE-4-18-756-1988 DE-4-18-758-1988 DE-4-18-759-1988 DE-4-34-1-1988 DE-4-34-3-1988 DE-4-34-5-1988 DE-4-34-11-1988 DE-4-34-12-1988 DE-4-34-15-1988 DE-4-34-16-1988 DE-4-34-19-1988 DE-4-34-20-1988 DE-4-34-30-1988 DE-4-34-31-1988 DE-4-41-1-1988 DE-4-41-2-1988 DE-4-41-4-1988 DE-4-41-5-1988 DE-4-41-9-1988 DE-4-41-10-1988 DE-4-41-12-1988
6 daughter colonies of queens mated at the mating station were tested:
DE-4-18-765-1989 DE-4-18-766-1989 DE-4-18-769-1989 DE-4-18-770-1989 DE-4-18-771-1989 DE-4-18-774-1989