53 daughter colonies of the 4a were tested:
DE-6-115-71-2021 DE-6-115-73-2021 DE-6-115-75-2021 DE-6-115-76-2021 DE-6-115-77-2021 DE-6-115-78-2021 DE-6-115-87-2021 DE-6-115-101-2020 DE-6-115-102-2020 DE-6-115-103-2020 DE-6-115-104-2020 DE-6-115-105-2020 DE-6-115-106-2020 DE-6-115-107-2020 DE-6-115-108-2020 DE-6-115-109-2020 DE-6-115-110-2020 DE-6-115-111-2020 DE-6-115-112-2020 DE-6-115-113-2020 DE-6-115-114-2020 DE-6-115-115-2020 DE-6-115-116-2020 DE-6-115-117-2020 DE-6-115-118-2020 DE-6-115-119-2020 DE-6-115-120-2020 DE-6-115-121-2020 DE-6-115-122-2020 DE-6-115-123-2020 DE-6-115-124-2020 DE-6-115-125-2020 DE-6-115-126-2020 DE-6-115-127-2020 DE-6-115-128-2020 DE-6-115-129-2020 DE-6-115-130-2020 DE-17-2-13-2020 DE-17-2-14-2020 DE-17-2-26-2020 DE-17-2-43-2020 DE-17-2-47-2020 DE-17-2-48-2020 DE-17-238-5-2020 DE-17-238-29-2020 DE-17-238-30-2020 DE-17-238-32-2020 DE-17-238-42-2020 DE-17-238-45-2020 DE-17-238-56-2020 DE-17-238-78-2020 DE-17-238-79-2020 DE-17-238-82-2020
374 tested queens were mated at the mating station:
DE-4-1-95-2021 DE-4-1-97-2021 DE-4-1-141-2021 DE-4-1-143-2021 DE-4-306-227-2021 DE-4-306-540-2021 DE-4-306-555-2021 DE-4-323-751-2021 DE-4-323-752-2021 DE-4-323-753-2021 DE-4-323-754-2021 DE-4-323-1402-2021 DE-4-323-1407-2021 DE-4-323-1408-2021 DE-4-323-1808-2021 DE-4-323-10310-2021 DE-4-323-10412-2021 DE-4-323-11001-2021 DE-4-323-11201-2021 DE-4-323-11305-2021 DE-4-323-11403-2021 DE-4-323-11404-2021 DE-4-323-11509-2021 DE-4-323-11604-2021 DE-4-323-11700-2021 DE-4-323-11707-2021 DE-4-323-11803-2021 DE-4-323-11901-2021 DE-4-323-11903-2021 DE-4-323-12507-2021 DE-4-323-12806-2021 DE-4-323-13205-2021 DE-4-323-13211-2021 DE-4-323-13309-2021 DE-4-323-13407-2021 DE-4-323-13610-2021 DE-4-323-13705-2021 DE-4-323-13903-2021 DE-4-323-14302-2021 DE-4-323-14511-2021 DE-4-323-14702-2021 DE-4-323-14703-2021 DE-4-323-15000-2021 DE-4-323-15010-2021 DE-4-323-15611-2021 DE-4-323-15906-2021 DE-4-323-16008-2021 DE-4-323-16309-2021 DE-4-323-16410-2021 DE-4-323-16509-2021 DE-4-323-17010-2021 DE-4-323-17206-2021 DE-4-323-17906-2021 DE-4-323-18007-2021 DE-4-323-18108-2021 DE-4-323-18403-2021 DE-4-323-18700-2021 DE-4-323-19010-2021 DE-4-323-19311-2021 DE-4-323-19903-2021 DE-4-323-22301-2021 DE-6-42-1-2021 DE-6-42-2-2021 DE-6-42-3-2021 DE-6-42-4-2021 DE-6-42-5-2021 DE-6-42-6-2021 DE-6-42-7-2021 DE-6-42-8-2021 DE-6-42-9-2021 DE-6-42-10-2021 DE-6-42-11-2021 DE-6-42-12-2021 DE-6-42-13-2021 DE-6-42-14-2021 DE-6-42-15-2021 DE-6-42-16-2021 DE-6-42-17-2021 DE-6-42-18-2021 DE-6-90-27-2021 DE-6-90-45-2021 DE-6-90-47-2021 DE-6-90-49-2021 DE-6-90-50-2021 DE-6-90-51-2021 DE-6-90-55-2021 DE-6-90-56-2021 DE-6-90-75-2021 DE-6-90-88-2021 DE-6-115-38-2021 DE-6-115-39-2021 DE-6-115-41-2021 DE-6-115-43-2021 DE-6-115-45-2021 DE-6-115-46-2021 DE-6-115-58-2021 DE-6-115-81-2021 DE-6-115-91-2021 DE-6-115-92-2021 DE-6-115-99-2021 DE-6-124-6-2021 DE-6-124-8-2021 DE-6-124-9-2021 DE-6-124-10-2021 DE-6-124-11-2021 DE-6-131-42-2021 DE-6-131-45-2021 DE-6-131-51-2021 DE-6-131-53-2021 DE-6-131-57-2021 DE-6-131-61-2021 DE-6-131-63-2021 DE-6-131-64-2021 DE-6-131-115-2021 DE-6-131-127-2021 DE-6-172-1-2021 DE-6-172-2-2021 DE-6-172-3-2021 DE-6-172-4-2021 DE-6-172-5-2021 DE-6-172-6-2021 DE-6-172-7-2021 DE-6-172-8-2021 DE-6-172-9-2021 DE-6-172-10-2021 DE-6-172-11-2021 DE-6-172-12-2021 DE-6-172-13-2021 DE-6-172-14-2021 DE-6-172-15-2021 DE-6-172-16-2021 DE-6-172-17-2021 DE-6-172-18-2021 DE-6-198-13-2021 DE-6-198-83-2021 DE-6-198-84-2021 DE-6-198-85-2021 DE-6-198-86-2021 DE-6-198-87-2021 DE-6-198-88-2021 DE-6-198-89-2021 DE-6-198-90-2021 DE-6-198-91-2021 DE-6-198-92-2021 DE-6-198-93-2021 DE-6-198-94-2021 DE-6-198-95-2021 DE-6-207-53-2021 DE-6-207-58-2021 DE-6-207-59-2021 DE-6-207-64-2021 DE-6-207-65-2021 DE-6-207-69-2021 DE-6-207-70-2021 DE-6-207-71-2021 DE-6-207-72-2021 DE-6-207-74-2021 DE-6-207-75-2021 DE-6-207-79-2021 DE-6-207-81-2021 DE-6-297-1-2021 DE-6-297-2-2021 DE-6-297-3-2021 DE-6-297-4-2021 DE-6-297-5-2021 DE-6-297-6-2021 DE-7-146-78-2021 DE-7-146-85-2021 DE-7-146-86-2021 DE-7-165-1253-2021 DE-7-165-1255-2021 DE-7-165-1260-2021 DE-7-165-1261-2021 DE-7-165-1265-2021 DE-7-165-1273-2021 DE-7-165-1275-2021 DE-7-165-1279-2021 DE-7-165-1280-2021 DE-7-165-1281-2021 DE-7-165-1282-2021 DE-7-165-1293-2021 DE-7-165-1294-2021 DE-7-165-1295-2021 DE-7-165-1296-2021 DE-7-165-1297-2021 DE-7-165-1298-2021 DE-7-165-1305-2021 DE-7-165-1309-2021 DE-7-165-1311-2021 DE-7-165-1315-2021 DE-7-165-1316-2021 DE-7-165-1317-2021 DE-7-176-35-2021 DE-7-176-75-2021 DE-7-179-73-2021 DE-7-179-74-2021 DE-7-179-75-2021 DE-7-179-76-2021 DE-7-179-77-2021 DE-7-179-78-2021 DE-7-179-79-2021 DE-7-179-80-2021 DE-7-179-82-2021 DE-7-179-83-2021 DE-7-179-84-2021 DE-7-179-85-2021 DE-7-179-86-2021 DE-7-179-88-2021 DE-7-179-89-2021 DE-7-179-90-2021 DE-9-10-49-2021 DE-9-10-52-2021 DE-9-21-20-2021 DE-9-21-21-2021 DE-10-30-17-2021 DE-10-30-18-2021 DE-10-30-23-2021 DE-10-30-24-2021 DE-10-30-25-2021 DE-10-30-26-2021 DE-10-30-29-2021 DE-11-1-21416-2021 DE-11-1-21430-2021 DE-11-1-21442-2021 DE-11-1-21446-2021 DE-11-1-21460-2021 DE-11-1-21488-2021 DE-11-139-3-2021 DE-11-139-28-2021 DE-11-139-74-2021 DE-11-139-80-2021 DE-11-139-104-2021 DE-11-139-105-2021 DE-11-139-121-2021 DE-11-139-131-2021 DE-11-139-133-2021 DE-11-139-143-2021 DE-11-139-145-2021 DE-11-161-26-2021 DE-11-189-3-2021 DE-14-21-1326-2021 DE-14-184-3-2021 DE-14-184-4-2021 DE-14-189-500-2021 DE-14-189-501-2021 DE-17-2-1-2021 DE-17-2-2-2021 DE-17-2-4-2021 DE-17-2-5-2021 DE-17-2-6-2021 DE-17-2-8-2021 DE-17-112-2-2021 DE-17-112-3-2021 DE-17-112-47-2021 DE-17-112-48-2021 DE-17-112-49-2021 DE-17-112-50-2021 DE-17-184-1-2021 DE-17-184-2-2021 DE-17-184-3-2021 DE-17-184-6-2021 DE-17-184-7-2021 DE-17-184-11-2021 DE-17-184-12-2021 DE-17-184-14-2021 DE-17-184-15-2021 DE-17-184-16-2021 DE-17-184-17-2021 DE-17-184-18-2021 DE-17-184-20-2021 DE-17-184-21-2021 DE-17-208-54-2021 DE-17-208-55-2021 DE-17-208-56-2021 DE-17-208-57-2021 DE-17-208-58-2021 DE-17-208-59-2021 DE-17-208-60-2021 DE-17-208-61-2021 DE-17-208-62-2021 DE-17-208-63-2021 DE-17-208-64-2021 DE-17-208-65-2021 DE-17-208-247-2021 DE-17-228-1-2021 DE-17-228-2-2021 DE-17-228-3-2021 DE-17-228-4-2021 DE-17-228-5-2021 DE-17-228-6-2021 DE-17-228-7-2021 DE-17-228-18-2021 DE-17-259-1-2021 DE-17-259-2-2021 DE-17-259-3-2021 DE-17-259-4-2021 DE-17-259-5-2021 DE-17-277-1-2021 DE-17-277-2-2021 DE-17-277-3-2021 DE-17-277-4-2021 DE-17-277-5-2021 DE-17-277-6-2021 DE-17-277-7-2021 DE-17-277-8-2021 DE-17-277-9-2021 DE-17-277-10-2021 DE-17-277-11-2021 DE-17-277-12-2021 DE-17-277-13-2021 DE-17-277-14-2021 DE-17-277-15-2021 DE-17-277-16-2021 DE-17-277-17-2021 DE-17-277-18-2021 DE-17-277-19-2021 DE-17-277-20-2021 DE-17-277-21-2021 DE-17-277-22-2021 DE-17-277-23-2021 DE-17-277-24-2021 DE-17-277-25-2021 DE-17-277-26-2021 DE-17-277-27-2021 DE-17-277-28-2021 DE-17-277-29-2021 DE-17-277-30-2021 DE-17-277-31-2021 DE-17-277-32-2021 DE-17-280-1-2021 DE-17-280-2-2021 DE-17-280-3-2021 DE-17-280-4-2021 DE-17-280-5-2021 DE-17-280-6-2021 DE-17-280-7-2021 DE-17-280-8-2021 DE-17-280-9-2021 DE-17-280-10-2021 DE-17-280-11-2021 DE-17-280-12-2021 DE-17-307-1-2021 DE-17-307-2-2021 DE-17-307-3-2021 DE-17-307-4-2021 DE-17-307-5-2021 DE-18-27-17-2021 DE-18-27-18-2021 DE-18-27-19-2021 DE-18-27-21-2021 DE-18-206-70-2021 DE-18-206-71-2021 DE-18-206-72-2021 DE-18-206-73-2021 DE-18-206-74-2021 DE-18-501-57-2021 DE-18-501-98-2021 DE-18-801-62-2021 DE-18-801-63-2021 DE-18-801-65-2021 DE-18-801-67-2021 DE-18-801-71-2021 DE-18-801-72-2021 DE-18-801-73-2021 DE-18-801-74-2021 DE-18-801-75-2021 DE-18-801-76-2021 AT-99-671-226-2021 NL-55-16-17-2021 NL-55-16-18-2021 NL-55-16-36-2021 NL-55-16-37-2021 NL-55-57-16-2021 NL-55-998-19-2021
408 daughter colonies of queens mated at the mating station were tested:
DE-2-363-13-2022 DE-2-363-16-2022 DE-2-363-17-2022 DE-2-613-3-2022 DE-2-613-4-2022 DE-2-613-12-2022 DE-2-613-18-2022 DE-2-613-19-2022 DE-2-613-36-2022 DE-2-613-47-2022 DE-2-613-49-2022 DE-2-613-50-2022 DE-2-613-57-2022 DE-2-613-60-2022 DE-2-613-65-2022 DE-2-613-87-2022 DE-2-613-91-2022 DE-2-749-87-2022 DE-2-749-94-2022 DE-2-749-95-2022 DE-4-306-1-2023 DE-4-306-2-2023 DE-4-306-5-2023 DE-4-306-8-2023 DE-4-306-10-2023 DE-4-306-16-2023 DE-4-306-19-2023 DE-4-306-24-2023 DE-4-306-25-2023 DE-4-306-34-2023 DE-4-306-35-2023 DE-4-306-42-2023 DE-4-306-43-2023 DE-4-306-47-2023 DE-4-306-79-2023 DE-4-306-83-2023 DE-4-323-103-2023 DE-4-323-111-2023 DE-4-323-112-2023 DE-4-323-113-2023 DE-4-323-114-2023 DE-4-323-127-2023 DE-4-323-152-2023 DE-4-323-154-2023 DE-4-323-155-2023 DE-4-323-156-2023 DE-4-323-157-2023 DE-4-323-159-2023 DE-4-323-161-2023 DE-4-323-162-2023 DE-4-323-165-2023 DE-4-323-167-2023 DE-4-323-168-2023 DE-4-323-171-2023 DE-4-323-175-2023 DE-4-323-176-2023 DE-4-323-177-2023 DE-4-323-178-2023 DE-4-323-183-2023 DE-4-323-185-2023 DE-4-323-191-2023 DE-4-323-194-2023 DE-4-323-198-2023 DE-6-1-285-2023 DE-6-1-288-2023 DE-6-1-291-2023 DE-6-1-292-2023 DE-6-1-701-2023 DE-6-1-747-2023 DE-6-1-751-2023 DE-6-1-752-2023 DE-6-1-755-2023 DE-6-1-757-2023 DE-6-1-759-2023 DE-6-1-763-2023 DE-6-1-765-2023 DE-6-1-771-2023 DE-6-1-774-2023 DE-6-36-1-2023 DE-6-36-2-2023 DE-6-36-3-2023 DE-6-36-4-2023 DE-6-36-5-2023 DE-6-36-6-2023 DE-6-36-7-2023 DE-6-36-8-2023 DE-6-36-9-2023 DE-6-36-10-2023 DE-6-36-11-2023 DE-6-36-12-2023 DE-6-36-13-2023 DE-6-36-14-2023 DE-6-36-15-2023 DE-6-36-16-2023 DE-6-36-17-2023 DE-6-36-18-2023 DE-6-36-19-2023 DE-6-36-20-2023 DE-6-36-21-2023 DE-6-36-22-2023 DE-6-36-23-2023 DE-6-39-1-2023 DE-6-39-2-2023 DE-6-39-3-2023 DE-6-39-4-2023 DE-6-39-5-2023 DE-6-39-6-2023 DE-6-39-7-2023 DE-6-39-8-2023 DE-6-39-9-2023 DE-6-39-10-2023 DE-6-39-11-2023 DE-6-39-12-2023 DE-6-39-13-2023 DE-6-39-14-2023 DE-6-39-15-2023 DE-6-39-16-2023 DE-6-39-17-2023 DE-6-90-21-2023 DE-6-90-23-2023 DE-6-90-24-2023 DE-6-90-26-2023 DE-6-90-28-2023 DE-6-90-29-2023 DE-6-90-31-2023 DE-6-90-36-2023 DE-6-90-39-2023 DE-6-90-45-2023 DE-6-90-47-2023 DE-6-90-48-2023 DE-6-90-55-2023 DE-6-90-57-2023 DE-6-90-58-2023 DE-6-90-59-2023 DE-6-90-63-2023 DE-6-90-65-2023 DE-6-90-82-2023 DE-6-90-85-2023 DE-6-90-86-2023 DE-6-90-87-2023 DE-6-90-88-2023 DE-6-90-98-2023 DE-6-115-307-2023 DE-6-131-70-2022 DE-6-172-13-2023 DE-6-172-14-2023 DE-6-172-15-2023 DE-6-172-16-2023 DE-6-172-17-2023 DE-6-172-18-2023 DE-6-172-19-2023 DE-6-172-20-2023 DE-6-172-21-2023 DE-6-172-22-2023 DE-6-172-23-2023 DE-6-172-24-2023 DE-6-172-25-2023 DE-6-172-26-2023 DE-6-172-27-2023 DE-6-172-28-2023 DE-6-172-29-2023 DE-6-172-30-2023 DE-6-172-31-2023 DE-6-172-32-2023 DE-6-172-33-2023 DE-6-172-34-2023 DE-6-172-35-2023 DE-6-297-1-2022 DE-6-297-2-2022 DE-6-297-3-2022 DE-6-297-4-2022 DE-6-297-5-2022 DE-6-297-6-2022 DE-6-297-7-2022 DE-6-297-8-2022 DE-6-297-9-2022 DE-6-301-3-2023 DE-6-301-5-2023 DE-6-301-6-2023 DE-6-301-8-2023 DE-6-301-9-2023 DE-6-301-10-2023 DE-6-301-11-2023 DE-6-301-12-2023 DE-6-301-13-2023 DE-6-301-15-2023 DE-6-301-16-2023 DE-6-301-17-2023 DE-6-301-18-2023 DE-6-301-25-2023 DE-6-319-37-2023 DE-6-319-58-2023 DE-6-322-1-2023 DE-6-322-2-2023 DE-6-322-3-2023 DE-6-322-4-2023 DE-6-322-5-2023 DE-6-322-6-2023 DE-6-322-8-2023 DE-6-322-11-2023 DE-6-322-12-2023 DE-6-322-13-2023 DE-6-322-14-2023 DE-6-322-54-2023 DE-6-322-55-2023 DE-6-322-56-2023 DE-6-322-57-2023 DE-6-322-58-2023 DE-6-322-59-2023 DE-6-323-3-2023 DE-6-323-4-2023 DE-6-323-15-2023 DE-6-323-21-2023 DE-6-324-71-2023 DE-6-324-72-2023 DE-6-324-81-2023 DE-6-324-82-2023 DE-6-324-85-2023 DE-6-324-86-2023 DE-6-324-87-2023 DE-6-324-88-2023 DE-6-324-89-2023 DE-6-324-91-2023 DE-6-324-92-2023 DE-6-324-95-2023 DE-6-324-96-2023 DE-6-324-97-2023 DE-6-324-98-2023 DE-7-146-3-2023 DE-7-146-8-2023 DE-7-146-11-2023 DE-7-146-15-2023 DE-7-146-31-2023 DE-7-146-2023-2023 DE-7-165-74-2023 DE-7-165-88-2023 DE-7-165-1582-2023 DE-7-165-1583-2023 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