4 daughter colonies of the 4a were tested:
AT-99-183-12-2011 AT-99-183-14-2009 AT-99-183-46-2009 AT-99-183-50-2011
27 tested queens were mated at the mating station:
AT-99-130-10-2011 AT-99-130-20-2011 AT-99-130-25-2011 AT-99-130-36-2011 AT-99-130-40-2011 AT-99-130-55-2011 AT-99-130-93-2011 AT-99-130-94-2011 AT-99-183-3-2011 AT-99-183-10-2011 AT-99-183-11-2011 AT-99-183-15-2011 AT-99-183-183-2011 AT-99-183-184-2011 AT-99-192-5-2011 AT-99-192-8-2011 AT-99-192-17-2011 AT-99-192-21-2011 AT-99-192-22-2011 AT-99-667-1-2011 AT-99-667-2-2011 AT-99-679-10-2011 AT-99-679-11-2011 AT-99-679-23-2011 AT-99-679-25-2011 AT-99-679-28-2011 AT-99-679-29-2011
15 daughter colonies of queens mated at the mating station were tested:
AT-99-130-2-2012 AT-99-667-5-2013 AT-99-667-11-2013 AT-99-679-20-2012 AT-99-679-21-2012 AT-99-679-22-2012 AT-99-679-48-2013 AT-99-679-49-2013 AT-99-679-50-2012 AT-99-679-52-2012 AT-99-679-54-2012 AT-99-679-55-2012 AT-99-679-62-2012 AT-99-679-64-2012 AT-99-679-65-2012