5 daughter colonies of the 4a were tested:
AT-99-689-16-2011 AT-99-689-17-2011 AT-99-689-19-2011 AT-99-689-20-2011 AT-99-689-21-2011
19 tested queens were mated at the mating station:
AT-99-160-81017-2012 AT-99-160-81025-2012 AT-99-160-91030-2012 AT-99-622-66855-2012 AT-99-622-66856-2012 AT-99-622-66857-2012 AT-99-622-66858-2012 AT-99-622-66859-2012 AT-99-622-66860-2012 AT-99-622-66861-2012 AT-99-622-66862-2012 AT-99-622-66863-2012 AT-99-622-66864-2012 AT-99-622-66866-2012 AT-99-640-81015-2012 AT-99-640-81020-2012 AT-99-640-81021-2012 AT-99-640-81046-2012 AT-99-640-81048-2012
19 daughter colonies of queens mated at the mating station were tested:
AT-99-622-66802-2014 AT-99-622-66803-2014 AT-99-622-66803-2015 AT-99-622-66804-2014 AT-99-622-66805-2014 AT-99-622-66805-2015 AT-99-622-66806-2014 AT-99-622-66806-2015 AT-99-622-66807-2014 AT-99-622-66807-2015 AT-99-622-66808-2015 AT-99-622-66809-2015 AT-99-622-66810-2015 AT-99-622-66811-2015 AT-99-622-66817-2014 AT-99-622-66819-2014 AT-99-622-66820-2014 AT-99-622-66821-2014 AT-99-622-66822-2014