17 daughter colonies of the 4a were tested:
DE-6-1-942-2016 DE-6-1-2010-2016 DE-6-1-2016-2016 DE-6-1-2017-2016 DE-6-1-2021-2016 DE-6-1-2022-2016 DE-6-1-2024-2016 DE-6-1-2027-2016 DE-6-1-2028-2016 DE-6-1-2033-2016 DE-6-1-2040-2016 DE-6-1-2041-2016 DE-6-1-2051-2016 DE-6-1-2052-2016 DE-6-1-2055-2016 DE-6-1-2056-2016 DE-6-1-2064-2016
67 tested queens were mated at the mating station:
DE-2-210-467-2017 DE-6-1-210-2017 DE-6-1-211-2017 DE-6-1-326-2017 DE-6-1-328-2017 DE-6-1-329-2017 DE-6-1-330-2017 DE-6-1-331-2017 DE-6-1-332-2017 DE-6-1-333-2017 DE-6-1-337-2017 DE-6-1-338-2017 DE-6-1-340-2017 DE-6-1-347-2017 DE-6-1-350-2017 DE-6-1-352-2017 DE-6-1-353-2017 DE-6-1-354-2017 DE-6-1-355-2017 DE-6-1-356-2017 DE-6-1-357-2017 DE-6-1-359-2017 DE-6-1-360-2017 DE-6-1-362-2017 DE-6-1-363-2017 DE-6-1-364-2017 DE-6-1-366-2017 DE-6-1-368-2017 DE-6-1-370-2017 DE-6-1-374-2017 DE-6-1-375-2017 DE-6-1-376-2017 DE-6-1-379-2017 DE-6-1-382-2017 DE-6-1-388-2017 DE-6-1-389-2017 DE-6-1-391-2017 DE-6-1-395-2017 DE-6-1-410-2017 DE-6-1-456-2017 DE-6-1-506-2017 DE-6-1-508-2017 DE-6-1-512-2017 DE-6-1-532-2017 DE-6-1-535-2017 DE-6-1-538-2017 DE-6-1-543-2017 DE-6-1-548-2017 DE-6-1-872-2017 DE-6-1-884-2017 DE-6-1-889-2017 DE-6-1-1022-2017 DE-6-1-1037-2017 DE-6-1-1511-2017 DE-6-1-2010-2017 DE-6-1-2012-2017 DE-6-1-2033-2017 DE-6-1-2042-2017 DE-6-1-2052-2017 DE-6-1-2083-2017 DE-6-1-2084-2017 DE-6-1-2086-2017 DE-6-1-2094-2017 DE-6-1-2098-2017 DE-6-1-2181-2017 DE-6-1-2183-2017 DE-6-1-2200-2017
83 daughter colonies of queens mated at the mating station were tested:
DE-6-1-115-2019 DE-6-1-301-2019 DE-6-1-326-2019 DE-6-1-334-2019 DE-6-1-342-2019 DE-6-1-349-2019 DE-6-1-351-2019 DE-6-1-420-2019 DE-6-1-494-2019 DE-6-1-513-2019 DE-6-1-906-2019 DE-6-1-911-2019 DE-6-1-920-2019 DE-6-1-924-2019 DE-6-1-1030-2019 DE-6-1-1031-2019 DE-6-1-1094-2019 DE-6-1-1122-2019 DE-6-1-1125-2019 DE-6-1-1129-2019 DE-6-1-1135-2019 DE-6-1-1157-2019 DE-6-1-1180-2019 DE-6-1-1211-2019 DE-6-1-1780-2019 DE-6-1-1811-2019 DE-6-1-1814-2019 DE-6-1-1828-2019 DE-6-1-1866-2019 DE-6-1-1883-2019 DE-6-1-1893-2019 DE-6-1-1900-2019 DE-6-1-1905-2019 DE-6-1-1925-2019 DE-6-1-1927-2019 DE-6-1-2067-2019 DE-6-1-2106-2019 DE-6-1-2108-2019 DE-6-1-2109-2019 DE-6-1-2202-2019 DE-6-1-2208-2019 DE-6-1-2248-2019 DE-6-1-2266-2019 DE-6-1-2324-2019 DE-6-1-2371-2019 DE-6-1-2411-2019 DE-6-1-2415-2019 DE-6-1-2416-2019 DE-6-1-2451-2019 DE-6-1-2467-2019 DE-6-1-2484-2019 DE-6-1-2604-2019 DE-6-1-2776-2019 DE-6-284-9-2019 DE-6-284-10-2019 DE-6-284-11-2019 DE-6-284-12-2019 DE-6-284-13-2019 DE-6-284-14-2019 DE-6-284-15-2019 DE-6-284-16-2019 DE-15-97-1-2019 DE-15-97-3-2018 DE-15-97-3-2019 DE-15-97-5-2018 DE-15-97-5-2019 DE-15-97-7-2018 DE-15-97-7-2019 DE-15-97-9-2018 DE-15-97-9-2019 DE-15-97-11-2019 DE-15-97-13-2019 DE-15-97-14-2019 DE-15-97-17-2019 DE-15-97-18-2019 DE-15-97-19-2019 DE-15-97-20-2019 DE-15-97-21-2019 DE-15-97-24-2019 DE-15-97-25-2019 DE-15-97-30-2019 DE-15-97-31-2019 DE-18-51-1480-2019